Fossil Crocs of 2022
For what its worth, its been a great year for fossil Pseudosuchians, many of which didn't get a lot of attention because people always assume its just "another crocodile". We'll go through them by the order of appearance in the fossil record, or in other words oldest to youngest. Better grab some popcorn and the drink of your choice, cause this is going to be a long one.
Mambawakale
Starting off is Mambawakale ruhuhu (Ruhuhu Basin Crocodile Ancestor in Kiswahili) from the middle Triassic Manda Beds of Tanzania. The fossils for this guy had been known since at least the 1960s, after which it was named Pallisteria angustimentum, a name that was never officially published however. It was a large animal, its skull 75 centimeters long attached to what was likely an animal similar in build to what we think of for "rauisuchians", given that it sits at the base of Paracrocodylomorpha. The below life reconstructoin was done by the fantastic Gabriel Ugueto with Steve Irwin inserted in post by me, tho its not rigorously scaled it gives a general idea of the animal's size.
We can then skip over the Jurassic (no new crocs from that period) and straight into the Cretaceous, specifically the Albian some 100 million years ago.
Yanjisuchus
Yanjisuchus longshanensis (Yanji City Crocodile from the Longshan Hill) was a paralligatorid, a group of Neosuchians I fully admitt I'm not too well aquainted with. Its fossils, which include a partial skull, vertebrae osteoderms, ribs, parts of the shoulder girdle and front limbs, were discovered in the Longjing Formation in what is now north-eastern China.
Confractosuchus
One I can decidedly talk more about is Confractosuchus sauroktonos (Lizard Eating Broken Crocodile) from the Australian Winton Formation, home to animals such as Isisfordia, Australovenator and Diamantinasaurus. The genus name refers to the fact that the skull is squashed to hell and back, but its the species name thats interesting. "Lizard eating" in this case refers to the fact that this 2.5 meter long basal Eusuchian was found with preserved stomach contents. Said stomach contents being the bones of a small ornithopod dinosaur as the press release art by Julius Csotonyi enthusiastically shows. The other image here not from the paper was drawn by Joschua Knüppe (@knuppitalism-with-ue ) showing a definitely more relaxed individual.
Eptalofosuchus
We did know about this guy for a bit longer as the pre-proof has been out since 2021, but lets include it anyways because I wasn't on Tumblr last year. Eptalofosuchus viridi (Green Seven Hills Crocodile) is not a crocodile in the strict sense but a Notosuchian, a cousin to the more typical Neosuchians. This animal is known from a single distinct lower jaw that belonged to a rather small creature. Among Notosuchians, Eptalofosuchus is most closely related to "advanced Notosuchians", specifically Sphagesauridae. We also know that it wasn't alone, as described alongside it are fossils of an unnamed baurusuchid (a large carnivore) and a peirosaurid, a more basal Notosuchian. Finally, both parts of the scientific name allude to the Uberaba Formation, specifically the nickname of Uberaba being “the city of the seven hills” and the green color of the sediments there. Life reconstruction of the little guy was done by the always incredible Júlia d'Oliveira.
Titanochampsa
Sticking to South America we have Titanochampsa iorii (Ior's Titan Crocodile). A significant find no doubt tho with an estimated size between 2.98–5.88 m (sizable I admitt) based on very scant remains perhaps a little bit hasty. Alas we only have the skull table of this animal, so the precise size can still vary as you can see. But it is still a fascinating one due to the fact that its the ONLY non-Notosuchian from the Bauru Group. Cretaceous South America is almost synonymous with these mostly terrestrial crocodyliforms but surprisingly lacking a diverse semi-aquatic croc fauna. Being at the very least a Neosuchian, possibly Eusuchian, Titanochampsa seems to have filled this niche. Given that the big terrestrial Baurusuchids had relatively weak bites and reached a length of at most 4 meters, this presents an interesting contrast with the strong bite force of the potentially larger semi-aquatic Titanochampsa. And would you look at that, press release art by Júlia once again.
Eurycephalosuchus
One final croc for the Mesozoic, Eurycephalosuchus gannanensis is an animal I covered very recently so I'll keep things brief. Stemming from the Maastrichtian Hekou Formation in south eastern China, Eurycephalosuchus was a very small species of orientalosuchine. Orientalosuchina is recovered as a group of early alligatoroids in the paper, tho talking to Adam Yates (who's croc phylogeny is my go-to) there is a chance they might be closer to crocodiloids. Regardless, Eurycephalosuchus was a small animal with large teeth towards the front of the skull, blunt teeth in the back and an overall very short face (tho there is slight compression so its not 1:1 like the fossil). Over on twitter people started comparing it to DnD Kobolds, leading to the below artwork by Manusuchus (@ mmujicam2000; full body of course highly speculative lmao)
Qianshanosuchus
Following the cataclysm that was KPG, crocs held on and in China we next meet Qianshanosuchus youngi (Young's Qianshan Basin Crocodile). It's another tricky one. While important as being the first crocodyloid from Paleocene China, the material, as you could guess based on the tiny size, is that of a juvenile (I now realize this being kinda ironic given the small but adult croc above). So the phylogeny is hard to nail down and gave different results. Still it could hold significance in our understanding how crocodyloids spread around the world and entered Europe after KPG. Below a quick comparison of the material with my hand.
Coming up a duo of two new species but not new genera, I also didn't really go deep into their publication as with some of the others so I'll try to stay brief
Maomingosuchus acutirostris
Maomingosuchus acutirostris (Acute Snouted Maoming Crocodile) is a new species of the already established genus Maomingosuchus from China. This new species however, differentiated amongst other things by having a pointed, not rounded, premaxilla stems from the Eocene of northern Vietnam. The skull is about 55 cm long giving us a medium sized tomistomines. Like Qianshanosuchus, M. acutirostris could have implications for the dispersal of crocodilians, in this case the spread of tomistomines from Europe to Asia, which the authors suggest happened thrice. That being said it should be noted that, between the two main ideas of what tomistomines are, the paper goes with them being a sister group to crocs and unrelated to modern gharials as Lee and Yates suggest (the later idea like I said I am more drawn towards).
Diplocynodon kochi
Another new species in a familiar genus, Diplocynodon kochi (Koch's Double Dog Tooth) is the latest in a long list of Diplocynodon species, a genus of alligatoroid very prominent in the Paleogene of Europe. This newest species from Transylvania is known from some well preserved skull material and was recovered as being one of the most derived members of its genus. With a size of 1.76 meters it was a medium sized Diplocynodon species and small by our modern standards. What is interesting is that the fossilw as recovered from shallow marine sediments. This is unusual in so far that alligatoroids are not especially known for their salt tolerance. Tho there's exceptions like Deinosuchus, alligators usually don't venture into salt water unlike crocodiles. Still, given the shallow water and the ability of large adult Mississippie Alligators to tolerate brackish water, it might have occasionally entered coastal waters. It is noted that the skull is in such good condition that it couldn't have been transported for a very long time.
Kinyang
Back to new genera we got Kinyang (derived from various Nilotic terms for crocodile), a genus of "giant" dwarf crocodile from the Miocene of Kenya. Now this guy doesn't just introduce one new species but two: Kinyang mabokoensis (Maboko Island Crocodile) and Kinyang tchernovi (Tchernov's Crocodile). Occuring in western and northern Kenya, Kinyang was a prime example over the prominence osteolaemines (dwarf crocodiles) had in the early Cenozoic prior to the takeover of crocodylines. Kinyang was surprisingly large and had a robust, blunt skull with several traits clearly setting it apart from any modern species. Still there is some clues that could be taken. Given the robust nature and more widened skull rather than V-shaped, its likely that it went after prey as big or even bigger than itself. They appear to have inhabited the shores of lakes located in open forests and woodlands. Alas this might have been their doom too. Osteolaemines are known to build nests made of foliage and with Africa growing increasingly dryer around this point in earths history, both change in prey and environment could have been factors that drove the dwarf crocodiles into the rainforests they inhabit today while the more dry resistant Crocodylus took over their rolle in the grasslands. The first image shows all three known skulls, the second is a quick composite of the cranium and mandible of K. mabokoensis I put together before the third image was published (ironic) by Christopher Brochu.
Almost done, only two more to go.
Sacacosuchus
On the other side of the world, specifically the Sacaco locality of Peru's Pisco Formation (among others) we have Sacacosuchus cordovai (Córdova's Sacaco Crocodile). A medium sized and longirostrine "tomistomine" gharial, here is where we come in conflict between the crocodylid and gavialoid model of what these guys are. Earlier with Maomingosuchus you had the idea that tomistomines were related to crocodylids, well here we have tomistomines being a paraphyletic group leading up to derived gharials. There is some significance to Sacacosuchus for seemingly showing that gharials entered South America on two separate occassions, but imo that's not the coolest thing about it. No instead I prefer to talk about the fact that its not just a marine gharial, living in the coastal waters of the Pacific, but that it coexisted with a second gharial showing perfectly how they specialise in different things. While the contemporary Piscogavialis was notably larger (about 7 meters) it had a very narrow and long snout, better adapted at catching fish. Sacacosuchus meanwhile was only 4.3 meters long, but with much more robust jaws, showing that it was much more of a generalist than its relative. This is well shown in the reconstruction by Javier "Canelita" Herbozo, tho it doesn't show the immense size difference. The third iamge shows Rodolfo Salas-Gismondi with the material and the fourth illustration by Gabriel Ugueto does not feature Sacaco, but the much more gracile Piscogavialis.
Hanyusuchus
The final one, Hanyusuchus sinensis (Han Yu's Crocodile from China) is the youngest crocodile described this year. How young? Well the OLDEST records of the species date to the Chinese Bronze Age, 4.000 BC, while the most recent records indicate that it went extinct in the 15th century. Hanyusuchus is a large tomistomine with a length between 5.43–6.19 meters and, if historical reports are to be believed, a big appetite. Stories tell of large crocodiles in Southern China attacking livestock, deer, boats and even killing humans while other reports suggest that these animals were used to fill moats and fed with prisoners. Despite appearing relatively narrow snouted, people often underestimate just how dangerous such animals could be. Even the modern false gharial has reports of it attacking and eating humans. Unsurprisingly, in addition to spawning some unique folklore (like crocodiles turning into tigers during autumn), this lead to humans attacking and killing these animals in retaliation. Fossils from the Bronze Age bear the clear marks of human attacks with cut marks focused around the animals head and the neck. Given the age of this material humans likely used bronze axes to kill and decapitate Hanyusuchus, drying their heads afterwards. Things didn't get better later in history. Han Yu, poet and politician and namesake of the genus, once demanded that the crocodiles leave the Han River delta before threatening to kill them. Later in history soldiers were sent to kill them in masses to avenge the death of a child. Eventually all of this, coupled with habitat loss, lead to the extinction of Hanyusuchus a mere 600 years ago. The below reconstructions were done by Hikaru Amemiya and Joschua Knüppe respectively.
And that is basically every crocodile officially published in 2022 (I think). Personally, I think its been a fantastic year for Pseudosuchian taxa, with some unique and fascinating additions.