All for One created Nezu.
Nezu states in the Anime and Manga that nobody knows what he is, so it's natural to assume that he's an amalgamation of sorts. Where else have we found amalgamations in My Hero Academia? The nomus.
Originally the origin of the nomus isn't known, until Hikoroshi reveals that the flying nomu had been one of Izuku's childhood bullies. At the time we see him during the stain arc he unrecognizable.
Now there are a multitude of theories on how nomus are made, during my observations they seem to be a mixture of animal-like and human traits. The process of creating nomu's could likely be taking a human and drastically modifying them with bio engineering, something that's generally looked down upon in the medical and animal ethnics world.
But AFO didn't just start from humans, he had to be sure of his processes and do multiple tests before the perfection of the nomu creating process. It's already be theorized and possibly confirmed that Kurogiri is a nomu, just not as far in the mutation process.
We've seen several instances of mutation processes all throughout multiple fandoms. One such instance, that is one of my favorites, is the Witcher series.
It's explained that Witchers are powerful because of the mutations they go through, Witchers are made specifically to be able to take down monsters and beasts that most of the human population cannot successfully do without manpower and knowledge.
Nomu's are created for a purpose, which is similar to the Witchers. However, Nomu's are more similar to the Kett from the Mass Effect series in the fact that the Kett are created for combat purposes. Once you reach a certain point in game, the Kett become more powerful, this showing that the game lore means that the Kett evolve.
For those who aren't aware, the Kett are created via mutations made to them. Sounds familiar.
So what does this have to do with Nezu?
What if Nezu was one of the original experiments, it makes sense to start with animals before moving to humans, and mice and rats are fairly easy to obtain.
All that's required is to try and put a quirk in one and modify their composition. All for One could've had several IQ quirks, it's possible that he could've been trying to create a nomu with the ability to think and act on its own.
This would've likely been before Tenko Shimura was ever even born, All For One needed a loyal pupil and what better way than to make one.
Obviously it reached the point where All for One could've thought Nezu was loyal to him, and Nezu used this to escape.
Nezu is also suspiciously close to Nomu, isn't it? The first and last letters stay the same, something that isn't common in Japanese names, now it's revealed that ultimately Nezu is a play of the Japanese word for rat(Nezumi) but Nezu can directly translate to Not Sleeping.
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