Let me preface this: this is the first time I've written anything about a show and I'd love to do that thing with run-on sentences that says, "don't take me too seriously" but that might get annoying to read.
A while ago I made a graphic series The Gang as Animals. I tried to pick stuff that had to do with the show a bit (Dee a bird, Dennis had a cat in the one episode... Mac and Poppins, Charlie and rats) as well as really fitting with their character. A while ago I had said Frank was a human, but it still wasn't an animal. I watched the episode where he's called "the warthog" but I couldn't buy it, or twist the wikipedia entry enough to make it fit for me.
Frank is really a spiritual being. The most spiritual on the show. Oh shit, did I lose you?? Hear me out!
Honestly, the only other character I think of that goes through anything spiritual is Dennis and the experience he gets out of murdering or... whatever he does.
One example of Frank as a sort of representation of the human condition is The Gang Gets Stranded. Frank is able to give himself into debauchery, abuse his kids, and only donates money to an animal charity so he can "make a mockery of their cause". But he still has a moment with that rabbit! He feels things, he connects with things. He's not a sociopath. Even as a someone who can connect and empathize he still wrecks shit. Human history is to the brim with this shit. Humans are intelligent and emotional but mankind as a whole loves destroying and conquering almost more than anything.
Frank desires both luxury and challenge. A normal apex predator is content with being at the top, but human beings have a history of not being satisfied. Frank took this to the extreem by starting from scratch and living with Charlie, but he still keeps his money so he can have access to anything he wants. He always needs a purpose and some goal to reach while crushing everyone below him he can. He finds satisfaction in hookers, drugs, and Charlie but he's still not erm "spiritually fulfilled" even with all the access to those things he wants. Frank has an ability to "think big" whereas the rest of the gang does not, with a bit of an exception to Dennis. I think Frank is really a good gluttonous representation of human beings as a whole. Complex and horrible.