My Choice: Dinosaur Nanotyrannus lethaeus, from Late Cretaceous, USA!
I was originally going to do some sort of stegosaur for my choice this year, since I unfortunately didn’t have room to include one on the list, but with the recent news about Nanotyrannus I realized I simply had to join the hype.
Nanotyrannus lancensis was first described in 1946 as a species of Gorgosaurus based on a single skull fossil, then re-examined and renamed in 1988 as its own species, Nanotyrannus. For years, the species was intensely controversial. As the skull belonged to (what was thought to be a) juvenile, many paleontologists (and paleofans) debated that Nanotyrannus was merely the juvenile of a Tyrannosaurus rex, not its own species. This debate was finally settled this year, when a new paper was published describing an adult specimen of Nanotyrannus lancensis. Exquisitely preserved, this animal had fewer vertebrae and more teeth than an adult T. rex, along with other anatomical differences that would make Nanotyrannus being a juvenile T. rex impossible. Not only that, but the discovery also shed light on another fossil, the “Jane” specimen, previously argued to be a Nanotyrannus but also thought to be a subadult T. rex. Jane was reassigned as a new species of subadult Nanotyrannus, N. lethaeus. Having been denied species-ship for so long, Nanotyrannus is now up to two.
Of the two species, N. lethaeus was larger, and mature individuals may have reached a body mass of about 1,200 kg (2,600 lb), while N. lancensis may have weighed nearly 704 kg (1,552 lb). They were mid-sized tyrannosauroids, combining characteristics of more basal Appalachian tyrannosauroids and more derived Laramidian tyrannosaurids. It was slender, with longer legs than other predatory theropods of the time, suggesting the niche of a pursuit predator. They would have been significantly faster than T. rex, with an estimated maximum running speed of 45.3–49.8 km/h (28.1–30.9 mph), nearly twice higher than that of adult T. rex with an estimated maximum running speed of 20.9–29.1 km/h (13.0–18.1 mph). Their arms and hands were also proportionally longer and larger than T. rex, suggesting they may have used them to manipulate prey more often than their giant, distant relatives. Jane’s skull has healed puncture wounds, likely the result of another Nanotyrannus bite, which seemed to have been common in larger theropods.
Both Nanotyrannus species lived in the Hell Creek Formation, which spans parts of modern-day Montana, South Dakota, North Dakota, and Wyoming, USA. It would have lived alongside its larger relative Tyrannosaurus, as a mesopredator, being preyed on by adult rexes but also ironically preying on juvenile rexes itself. Nanotyrannus’ long legs would have been very helpful for allowing it to escape this apex predator. In Hell Creek, Nanotyrannus would have lived alongside many of the most famous North American dinosaurs. These included other theropods like the dromaeosaurs Acheroraptor and (possibly) Dakotaraptor, the ornithomimids Struthiomimus and Ornithomimus, the alvarezsaur Trierarchuncus, the caenagnathids Anzu and Eoneophron, and the avialans Avisaurus, Brodavis, Magnusavis, and Potamornis. The most common dinosaur of this formation was of course Triceratops, with two species, but other ceratopsians lived here as well, including Torosaurus, Tatankaceratops, and Leptoceratops. Other ornithischians Nanotyrannus could have come across include the giant hadrosaur Edmontosaurus annectens, the small Thescelosaurus, the pachycephalosaurs Pachycephalosaurus, Platytholus, and Sphaerotholus, and the ankylosaurs Ankylosaurus and Denversaurus. Other animals would have included the crocodylomorphs Borealosuchus, Brachychampsa, and Thoracosaurus, turtles like Axestemys, Aspideretoides, and Neurankylus, the choristodere Champsosaurus, and a variety of mammals. Nanotyrannus lethaeus was collected alongside the vertebrae of the azhdarchid Infernodrakon (which would have been just the right size for the mid-size tyrannosauroid to grab at neck height).
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