Multituberculate Earth: Paleocene
This where the timeline diverges.
The first five million years of the Paleocene occur as in our timeline. After a slow recovery multituberculates explode in diversity, dominating the mammal faunas of the north hemisphere. Meanwhile, in the southern continents, gondwanatheres and dryolestoids do the same. Therians radiate in the background, placentals in particularly producing a myriad of forms.
But while in our timeline multituberculates rapidly decline at the end of the Danian (circa 61 million years) and therians take hold, this doesn’t happen here. Instead, multituberculates continue their trajectory as dominant land mammals, while therians decline in their stead. Whatever caused the decline of multituberculates in our timeline doesn’t affect them here; it instead cuts the branches of the therian family tree, and by the end their brief period of exploration comes to an end.
The climate is temperate, warming as the era goes on. Flowering plants dominate the scene, but not as much as in our world, there being also a wealth of conifers, ferns, horsetails and ginkgos among others (this could explain why multituberculates have not declined, but fails to explain why therians did…) In these conditions various groups have prospered in the post-Cretaceous world, including the sole surviving dinosaurs, birds, as well as crocodylomorphs, turtles, amphibians and the mysterious choristodere reptiles. Squamates aside from snakes were strongly impacted by the KT event, and it took them over ten million years to recover in our timeline; this recovery does not happen here, making them join therians on the path to damnation.
In the north hemisphere, cimolodont multituberculates are the dominant mammals. Kogaionids make a brief tour throught Europe, but increasing competition from recently-arrived North American and Asian clades force them back to the isolation of Balkanatolia. Taeniolabidoids dominate herbivorous guilds, taeniolabidids as large sized herbivores across the northern continents and lambdopsalids as a myriad of digging, running and even hopping forms in Asia. Ptilodontoideans diversify across the canopies, and go on to produce the first largest multituberculate carnivores. Microcosmodontids remain largely small sized insectivores, but already vie for large sized predatory roles in some environments. Meniscoessids, inversely, occupy herbivorous roles and don’t diversify as much. Djadochtatheroideans, which evolved in Asia’s desert environments during the Cretaceous, have a hard time ajdusting to a greener world, but still succeed as the mostly predatory eucosmodontids and herbivorous boffiids.
In the trifecta of South America, Antarctica and Australia, gondwanatheres and dryolestoids continue the reign they enjoyed in the Cretaceous. The former are mostly herbivorous, while the latter fulfill all manner of roles from sengi-like runners to large carnivores to long-necked leaf-eaters that earn the title of largest land animal… for now at least. Curiously, some northern mammal groups penetrate into South America from North America much as in our world, but unlike our timeline marsupials are not the ones leaving a lasting legacy. And unlike our timeline, ferugliotheriid gondwanatheres invade the north successfully.
Africa stands as an isolated oddity. Here, a sole lineage of gondwanatheres, the galulatheriids, dominate herbivorous niches, while kogaionids diversify as carnivores and insectivores; whereas the latter have been here all along since the Cretaceous or are recent immigrants from Europe and/or Balkanatolia is unknown. Also weird are Madagascar and India, two isolated landmasses bearing a similar gondwanathere/kogaionid fauna, though the former are composed of sudamericids and adalatheriids instead. India is marching northwards at great speeds, and soon contact between the two faunas will begin…
Example Site: Santa Lucia Formation
Example Site: Khashat Formation