Triassic Double Feature
Just weeks ago I remissed the lack of new croc taxa, seems I spoke to soon because they are being pumped out like crazy right now. For simplicity, I will cover two of the recent sorta-crocs together as neither are super extensive and they match in overall time.
These two new genera are the aetosaur Kryphioparma and the phytosaur Jupijkam. Pictured below the fossils of both with reconstructions of close relatives (art by Brian Engh and Gabriel Ugueto).
I'll start with Kryphioparma, which I wager is the less interesting of the two. Kryphioparma was an aetosaur, which are effectively early pseudosuchians that evolved a body type very similar to what nodosaurs did later. Heavily armored, sometimes with prominent spikes and herbivorous.
Kryphioparma is only known from four isolated and incomplete osteoderms, and though these are actually diagnostic and highly distinct, it does mean there's not super much to say. Hell, the scientific name literally means "mysterious shield" in reference to how little we know.
Regardless, scientists did determine two things. 1) It's a typothoracine aetosaur, narrowing down its placement to one of the two main branches. This means its closer related to Typothorax (with its armored cloaca) than to Desmatosuchus (with its shoulder spikes). 2) The second thing we know is that it wasn't alone. No, the localities that yielded its bones (Placerias Quarry and Thunderstorm Ridge) actually preserve a highly diverse aetosaur fauna, including Calypsosuchus, Tecovasuchus and two species of Desmatosuchus. All images by Jeff Martz.
Now the arguably more exciting find is that of Jupijkam, a type of phytosaur, which are archosaurs superficially resembling today's crocodiles. Now when I grew up, phytosaurs used to be considered to be entirely unrelated to crocs, being a type of archosaur believed to have diverged prior to the bird-croc split. However, it would appear that recent studies suggest that they could actually be true croc-line archosaurs, potentially being the earliest diverging group of Pseudosuchian.
Jupijkam is from the Rhaetian-Norian Blomidon Formation of Novia Scotia, Canada. This not only makes it one of the youngest, but also one of the northern-most known phytosaurs to date. It's scientific name, Jupijkam, is actually derived from the name given by the local Mi’kmaq people to their version of the horned serpent.
How Jupijkam is related to other phytosaurs is a bit wonky. Now generally, its recovered as a mystriosuchine, which isn't exactly a surprise given that most phytosaurs fall into this category. It's placement however shifts ever so slightly depending on the precise methods and characters used. 2/4 times it was found to be most closely related to Rutiodon (the phytosaur shown at the very start), once alongside the Indian Volcanosuchus and once to its exclusion. One tree simply results in a big polytomy which isn't really helpful, and one time it was found as a much more derived form related to Mystriosuchus. Whatever the case, additional finds both of other phytosaurs and Jupijkam specifically might clarify this in the future. Currently however, it seems that this form is not related to all the other American species of its time, suggesting it held out till the late Triassic independently. Which is pretty neat. Below you can see a comparisson between Jupijkam and some other slender-snouted phytosaurs, courtesy of Brownstein 2023. A is Jupijkam, B is Rutiodon and C is Machaeroprosopus
Really that wraps things up already, two new genera, both Triassic, both (potentially) early Pseudosuchians. A little bit out of my usual focus but very interesting none the less. Definitely gotta make another post soon since they just dropped yet another new one (a metriorhynchid), but I gotta read that paper first. Speaking of which A new aetosaur (Archosauria: Pseudosuchia) from the upper Blue Mesa Member (Adamanian: Early–Mid Norian) of the Late Triassic Chinle Formation, northern Arizona, USA, and a review of the paratypothoracin Tecovasuchus across the southwestern USA (escholarship.org) A late-surviving phytosaur from the northern Atlantic rift reveals climate constraints on Triassic reptile biogeography | BMC Ecology and Evolution | Full Text (biomedcentral.com) plus the respective Wikipedia pages Kryphioparma - Wikipedia Jupijkam - Wikipedia