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Fossil Novembirb: Day 14 - Lost in the Woods

As the global climate was cooling during the Oligocene epoch, tropical rainforests started to shift towards their current distribution in the tropics, and open landscapes such as scrubland was spreading. But that doesn't mean forests were disappearing. In places like Central and Eastern Europe, forests shifted from tropical to temperate. And in these shifting landscapes, birds continued to adapt and diversify.

Turnipax: One of the earliest known members of a relatively obscure bird groups called buttonquails. Despite their quail-like appearance and lifestyle, buttonquails are shorebirds related to gulls and sandpipers.

Palaeotodus: This small insectivore is one of the first known members of the todies, a bird group nowadays restricted to the West Indies.

Primotrogon: An early member of the trogons which had a very different shape compared to its modern relatives, including long wings, a short tail, and relatively small eyes.

Eurotrochilus: The first known true hummingbird, a group now endemic to the Americas, but first appeared in Europe.

Aviraptor: A thrush-sized true hawk with long thin legs and sharp talons, this was a prime predator of small forest birds in the Oligocene.

Wieslochia: One of the earliest known passeriformes, a group that likely evolved in Australia,parts of it's skeleton resemble certain South American passeriforms, like cotingas.

Rupelramphastoides: As the earliest known ramphastid, it was closely related to the barbets as well as toucans, but it bore a closer resemblance to the former.

Oligocolius: A bizarre relative of modern mousebirds that had an appetite for seeds, as the holotype was preserved with seeds in its crop and gullet. It also had a slightly parrot-like beak.

Laurillardia: A distant relative of hoopoes and hornbills, this long winged and long tailed bird used its sharp beak to catch insects.

Rupelornis: A close relative of albatrosses, these seabirds had delicate beaks and long legs. They probably had a similar lifestyle to storm petrels, which carefully pick food from the surface of the water.

#Fossil Novembird #Novembird #Dinovember #birblr #palaeoblr #Birds #Dinosaurs #Cenozoic Birds #Turnipax #Palaeotodus #Primotrogon #Eurotrochilus #Aviraptor #Wieslochia #Rupelramphastoides #Oligocolius #Laurillardia #Rupelornis

Rupelramphastoides knopfi

By Ripley Cook

Etymology: Barbet from the Rupelian

First Described By: Mayr, 2005

Classification: Dinosauromorpha, Dinosauriformes, Dracohors, Dinosauria, Saurischia, Eusaurischia, Theropoda, Neotheropoda, Averostra, Tetanurae, Orionides, Avetheropoda, Coelurosauria, Tyrannoraptora, Maniraptoromorpha, Maniraptoriformes, Maniraptora, Pennaraptora, Paraves, Eumaniraptora, Averaptora, Avialae, Euavialae, Avebrevicauda, Pygostaylia, Ornithothoraces, Euornithes, Ornithuromorpha, Ornithurae, Neornithes, Neognathae, Neoaves, Inopinaves, Telluraves, Afroaves, Coraciimorphae, Cavitaves, Eucavitaves, Picocoraciae, Picodynastornithes, Piciformes, Pici, Ramphastides?

Status: Extinct

Time and Place: Between 31 and 30 million years ago, in the Rupelian of the Oligocene of the Paleogene

Rupelramphastoides is known from the Bott-Eder Grube-Unterfeld Quarry of Baden-Württemberg, Germany

Physical Description: Rupelramphastoides is the oldest known barbet-like bird - the barbets being a group of tree-dwelling birds that includes the iconic Toucans, though most barbest don’t have nearly as impressive beaks. This bird resembled its modern relatives in a lot of ways, which is notable given how old it is (still in the Paleogene, aka the first period of the time of modern birds). However, it is also one of the smallest members of the group overall, with very long and slender bones in its foot like modern toucans. Still, it had a small beak - more like non-toucan barbets - and a much smaller size than the living toucans. It had long and narrow arm bones but shorter fingers, giving the wings a more squat appearance at the ends. In general, its legs were slender, as were its toes, so perhaps Rupelramphastoides represents a transitional form between most barbets and the very weird toucans, though this is just conjecture. It probably wouldn’t have gotten much longer than 9 centimeters in overall body length, and it had a small head with a robust triangular beak.

Diet: It is uncertain what Rupelramphastoides ate, but like living barbets and toucans it probably mainly ate fruit, with mainly some supplementing of its diet with insects.

Behavior: Rupelramphastoides would have probably spent most of its time in the trees, hopping around and walking from branch to branch in search of food. It then probably would have stretched about to try and get food, using its beak to chomp into berries and other fruits. It could also use its long and skinny legs to stand up tall and reach food that it couldn’t otherwise. It probably lived in small flocks rather than big ones, and took care of their young. They also probably didn’t migrate. Like other members of the Pici, it was probably at least somewhat brightly colored, and used color in display to one another.

By Scott Reid

Ecosystem: Rupelramphastoides lived in a coastal/bay area, filled with estuaries and streams leading out into the ocean. This was a very lush habitat, filled to the brim with plants from algae ferns and cycads; to conifers, palms, roses, asterids, beeches, oaks, and cypresses. This dense coastal forest transitioned into water-based plants, and its possible there was some sort of a coastal swamp (a proto-mangrove swamp, even?) in the area. This is fascinating as this habitat existed just at the time of the global rainforest collapse, and the transition of the world into more varied and arid habitats - clearly, this particular coastal swamp was not hit by this event. Dozens of kinds of fish lived here, including things like trumpetfish, boarfishes, eels, ladyfish, sea robins, rockfish, shrimpfish, pipefish, sailfin moonfishes, halfbeaks, sea breams, weevers, and so many others that I just have to stop now. There were sharks and rays present as well, though not in as much diversity as the ray-finned fish. Insects such as beetles, flies, and butterflies were present, as well as spiders, crustaceans, snails, slugs, clams, oysters, and other invertebrates. Plenty of turtles were also present. Mammals were rarer in this habitat, but included extinct relatives of modern sea cows and bats; and a predatory Hyaenodont named Apterodon. Many different kinds of dinosaurs lived here alongside Rupelramphastoides, including the early hummingbird Eurotrochilus, the buttonquail Turnipax, the tody Palaeotodus, the (rare example of a) fossilized songbird Wieslochia, the mousebird Oligocolius, the trogon Primotrogon, the loon Colymboides, and the seabird Rupelornis that vaguely resembled modern albatrosses and petrels. In short, you could see a wide variety of ocean-going and tree-dwelling birds, as one would expect in a forested esturary area, and Rupelramphastoides probably mostly had to worry about predation from they Hyaenodonts in the area, rather than other birds.

Other: The exact phylogenetic affinities of Rupelramphastoides are very murkey, because a phylogenetic analysis has never been done on this dinosaur. Morphologically speaking it seems to be somewhere in the Barbet-Toucan clade, with similarities to both non-toucan barbets and the toucans, but it also has traits of other members of Pici such as the honeyguides, piculets, and woodpeckers, and so it can’t really be confirmed as a member of Ramphastides for now.

~ By Meig Dickson

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