The process of Photosynthesis Part 1
The process of photosynthesis is one of the key factor in the success of life on planet earth. It is the primary reason plants are autotrophic (more specifically; phototrophic), as glucose, their primary food source, is one of the products. Most people are familiar with the basic concept; plants convert light energy into storable carbohydrates (glucose), as well as fix carbon into plant tissue. But the intricacies of how plants achieve this are fascinating.
Photosynthesis occurs in the chloroplasts of mesophyll cells, which are found in mesophyll tissues which can be found on the shoots and primarily the leaves of plants. The basic stoichiometric equation for photosynthesis:
Light energy
6CO2 + 12H2O + Energy --> C6H12O6 + 6O2 + 6 H2O
Chloroplasts
The sugars are stored, and eventually used to generate ATP in respiration, which is ultimately used in biosynthesis within the plant.
There are two major pathways within photosynthesis; light, and dark reactions. The “light reactions” capture light energy and convert it into chemical energy within reaction centers. Light is trapped by pigments (chlorophyll a and b and carotenoids) and funneled by passing the resonance energy cell by cell “downhill” toward the reaction center. Water is also split during the light reactions (the mechanism of this is poorly understood) to release elections (e-), H+, and O2. The electron transfer then produces ATP from ADP and Pi, as well as NADPH from NADP and H+. Those products then feed into the calvin cycle (ATP and NADPH are high energy chemicals used in dark reactions as well). The light reaction is shown.
H2O + NADP+ + ADP + Pi + light → NADPH + H+ + ATP + ½ O2
The dark reactions convert chemical energy into sugars and require more ATP then NADPH, which is why a continuous flow of e- are needed. Ribulose bisphoshate carboxylase/oxygenase or Rubisco is an enzyme that catalyzes the dark reaction, and may be the most abundant protein on planet earth. A simplified equation for the dark reaction is shown
ATP + NADPH + CO2 ---> ADP + NADP + glucose
Each of these photosynthetic pathways is critical to the environment and shape how it functions today. The light reactions have allowed plants to dominate and characterize our environment by exploiting an abundant and novel resource (sun), and the dark reactions fix atmospheric carbon into plant tissue (a process especially important today). A more complex view of photosynthesis could be explained in a 2 hour power point lecture, including the “Z” scheme, photorespiration, and the proton pump which can be found in the “further reading”.
-Greg Aegis
Further Reading
http://faculty.clintoncc.suny.edu/faculty/michael.gregory/files/bio%20101/bio%20101%20lectures/photosynthesis/photosyn.htm
http://www.emc.maricopa.edu/faculty/farabee/biobk/biobookps.html
Photo Credit
http://www.pxlshots.com/contest/288/leaves.html