*Crocodile Dundee voice* Mate, that’s not Gustave:
And he is the PERFECT CROCODILE. He is the perfect example of what I mean when I talk about (as I do) how the morphology of extremely large crocodiles adapts to the changing physics of their bite.
This is a typical adult Nile Crocodile:
And THIS is a god among his kind:
This is it, folks. The Final Form. THIS is what peak performance looks like.
Crocodiles and physics have an interesting relationship. Crocodiles have, by a CONSIDERABLE MARGIN, the strongest bite of any animal on Earth. EVER. Scaled up estimates (based on Nile and Saltwater crocodiles) give the extinct Deinosuchus an estimated bite force MORE THAN DOUBLE the recently updated Tyrannosaurus bite estimates. Living crocodiles have bite forces measured in the range of 5000 pounds per square inch, for an individual around 15-16 feet. It is estimated that modern crocodiles in the range of 18-20 feet would have bit forces around 7-8000 psi or more.
Because a crocodile’s skull is only designed to handle so much pressure. Go beyond that limit and the force of impact when those jaws snap shut could literally shatter their own skulls.
But evolution has spent hundreds of millions of years perfecting crocodiles, so PHYSICS ISN’T GOING TO STOP THEM. What ends up happening in the skulls of these extremely large crocodiles is they will increase dramatically in mass to compensate for the increased forces. A crocodile’s skull is almost exclusively solid bone, with only minimal space for nasal passages, a surprisingly advanced brain, and some slightly porous looking framework that helps the bone distribute the force over a larger area. The effect is by far the most pronounced in Nile crocodiles, which most regularly feed on larger prey and need to make use of all that power.
Both of those are Nile crocodile skulls (or rather, replicas thereof).
And just for fun, here are the skulls of completely different (and very extinct species), Deinosuchus:
The bigger the crocodile (within a given species), the more massive the skull needs to be to compensate for that UNBELIEVABLE bit pressure. This is one way to see from a distance whether you are looking at a normal sized crocodile:
and a truly extraordinary individual:
One of the things about Gustave that’s so impressive is how healthy his teeth look. A lot of large crocodiles, in their old age, have very worn down and often missing teeth. They do replace them many times over a lifetime, but when they get very old this slows down. Gustave, at least in every picture taken of him, had teeth that were in very good condition.
Even crocodiles much smaller than Gustave’s reported size (probably similar in size to Dominator or Lolong) tend to have smaller or more worn teeth:
than the pinnacle of his kind: