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Born of Primus himself, these Avatars of His Will were created to guide and prosper the Planet that was their Creator and God.
The First were the Twins; Solus and Prima. While Prima forges the laws and ideals, Solus created foundations and forged artifacts that would help the future generations.
The Second of these beings was Onyx. His duty was to tend to the natural flora and fauna that came with his creation. These cybernetic lifeforms mimic our own, but with stark differences. Onyx himself was built similar to the animals that inhabit their world.
Third and Fourth were Alpha Trion and Vector Sigma. Guardians of Space and Time. While they both took care of recording every act the Grand 13 did– every gain and void, what they said and would do– Both of these wise mechs protected the flow of their world. Alpha Trion could manipulate the space around him; He could form images using metallic particles, and his alt mode was a powerful cyber-puma. Vector Sigma, on the other hand, was the keeper of time; his knowledge extended beyond the present and the past, making him a perfect scribe, until he transformed himself into a powerful entity, one that held all the knowledge of the Grand 13, so should future generations need it, he could grant it to them by way of a Key…
The rest followed after Vector Sigma transformed.
The Grand 13 could not be left alone, so Primus created his true avatar; a meager mech named Rung. Unassuming to everyone else but the 13. He was their mentor, their guide, and caretaker. He watched as Megatronus became possessed by a darkness so ancient, none of them knew what it was. He killed many of his brethren, even bots that weren't part of The Order. Solus herself, the conjunx of Megatronus, dealt the final blow, which cleansed him of his possession, with his final words, he whispered “I'm sorry…- I love you…”
Solus mourned. And she stopped creating. Foundations crumbled under her distress, and the Grand 13 were reduced to just Solus, Alpha Trion, Nova, and Quintus, and they couldn't console their grieving sister. Solus's spark break eventually caused her an early end, not that she did it voluntarily… Her spark couldn't handle the weight of what she'd done. Poor Rung couldn't help her, and went into hiding.
Around the time of Megatronus's fall, Quintus had created the Quintessons, to help in the fight against him. But he, along with his creations, were driven off the planet… But Quintus disappeared sometime after, not even the Quintessons knew where he went… Or so they said.
Alpha Trion found himself alone when Nova went off to find Rung, and he never returned. Alpha Trion lived a long time, up to the rise of a new council, and the intentions of their hierarchy. While Alpha Trion supported the potential, he didn't agree with how Civilians treated Warframes, especially after Energon stopped flowing following the fall. Many Magnus's, the highest influence of Cybertron, Alpha Trion guided. Subtly, he tried to obliterate the system that oppressed the Warframes. Unfortunately, he was unsuccessful, and Megatron arose as the leader of the Decepticons… And we all know what events followed after that.